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Chewing and Swallowing

Author: Sophia

what's covered
In this lesson, you will learn about the elements of chewing and swallowing as well as their roles in digestion. Specifically, this lesson will cover:

Table of Contents

1. The Mouth

The cheeks, tongue, and palate frame the mouth, which is also called the oral cavity (or buccal cavity).

At the entrance to the mouth are the lips, or labia (singular = labium). Their outer covering is skin, which transitions to a mucous membrane inside the mouth.

did you know
Lips are very vascular with a thin layer of keratin—hence, the reason they are "red." They also have a huge representation on the brain’s cerebral cortex, which probably explains the human fascination with kissing!

The lips cover the orbicularis oris muscle, which regulates what comes in and goes out of the mouth. The labial frenulum is a midline fold of mucous membrane that attaches the inner surface of each lip to the gum. The cheeks make up the oral cavity’s sidewalls. While their outer covering is skin, their inner covering is mucous membrane. This membrane is made up of non-keratinized, stratified squamous epithelium. Between the skin and mucous membranes are connective tissue and buccinator muscles.

try it
Directions: The next time you eat some food, notice how the buccinator muscles in your cheeks and the orbicularis oris muscle in your lips contract, helping you keep the food from falling out of your mouth. Additionally, notice how these muscles work when you are speaking.

The pocket-like part of the mouth that is framed on the inside by the gums and teeth, and on the outside by the cheeks and lips, is called the oral vestibule. Moving farther into the mouth, the opening between the oral cavity and throat (oropharynx) is called the fauces (like the kitchen "faucet"). The main open area of the mouth, or oral cavity proper, runs from the gums and teeth to the fauces.

When you are chewing, you probably do not find it difficult to breathe simultaneously. The next time you have food in your mouth, notice how the arched shape of the roof of your mouth allows you to handle both digestion and respiration at the same time. This arch is called the palate. The anterior (front) region of the palate serves as a wall (or septum) between the oral and nasal cavities as well as a rigid shelf against which the tongue can push food. It is created by the maxillary and palatine bones of the skull and, given its bony structure, is known as the hard palate.

If you run your tongue along the roof of your mouth, you’ll notice that the hard palate ends in the posterior oral cavity, and the tissue becomes fleshier. This part of the palate, known as the soft palate, is composed mainly of skeletal muscle. You can therefore manipulate, subconsciously, the soft palate—for instance, to yawn, swallow, or sing.

Mouth—The mouth includes the lips, tongue, palate, gums, and teeth.

A fleshy bead of tissue called the uvula drops down from the center of the posterior (back) edge of the soft palate. Although some have suggested that the uvula is a vestigial organ (an unused organ that does not currently have function), it serves an important purpose. When you swallow, the soft palate and uvula move upward, helping to keep foods and liquid from entering the nasal cavity. Unfortunately, it can also contribute to the sound produced by snoring. Two muscular folds extend downward from the soft palate, on either side of the uvula. Toward the front, the palatoglossal arch lies next to the base of the tongue; behind it, the palatopharyngeal arch forms the superior and lateral margins of the fauces.

Between these two arches are the palatine tonsils, which are clusters of lymphoid tissue that protect the pharynx. The lingual tonsils are located at the base of the tongue.

The digestive functions of the mouth are summarized in the table below, and you will learn more about some of these structures throughout the rest of this lesson.

Digestive Functions of the Mouth
Structure Action Outcome
Lips and cheeks Confine food between teeth
  • Food is chewed evenly during mastication
Salivary glands Secrete saliva
  • Moisten and lubricate the lining of the mouth and pharynx
  • Moisten, soften, and dissolve food
  • Clean the mouth and teeth
  • Salivary amylase breaks down starch
Tongue’s extrinsic muscles Move tongue sideways, and in and out
  • Manipulate food for chewing
  • Shape food into a bolus
  • Manipulate food for swallowing
Tongue’s intrinsic muscles Change tongue shape
  • Manipulate food for swallowing
Taste buds Sense food in mouth and sense taste
  • Nerve impulses from taste buds are conducted to salivary nuclei in the brain stem and then to salivary glands, stimulating saliva secretion
Lingual glands Secrete lingual lipase
  • Activated in the stomach
  • Break down triglycerides into fatty acids and diglycerides
Teeth Shred and crush food
  • Break down solid food into smaller particles for deglutition (swallowing)

terms to know
Oral Cavity
Referred to as the cheeks, tongue, and palate which frame the mouth.
Labia
The lips.
Oral Vestibule
The part of the mouth bounded externally by the cheeks and lips, and internally by the gums and teeth.
Hard Palate
The bony structure that forms the roof of the mouth and floor of the nasal cavity, formed by the palatine process of the maxillary bones and the horizontal plate of the palatine bones.
Soft Palate
The posterior region of the bottom portion of the nasal cavity that consists of skeletal muscle.

1a. The Teeth

The teeth, or dentes (singular = dens), are organs similar to bones that you use to tear, grind, and otherwise mechanically break down food.

During the course of your lifetime, you have two sets of teeth (one set of teeth is a dentition). Your 20 deciduous teeth, or baby teeth, first begin to appear at about 6 months of age. Between approximately age 6 and 12, these teeth are replaced by 32 permanent teeth. Moving from the center of the mouth toward the side, these are as follows:

  • The eight incisors, four top and four bottom, are the sharp front teeth you use for biting into food.
  • The four cuspids (or canines) flank the incisors and have a pointed edge (cusp) to tear up food. These fang-like teeth are superb for piercing tough or fleshy foods.
  • Posterior to the cuspids are the eight premolars (or bicuspids), which have an overall flatter shape with two rounded cusps useful for mashing foods.
  • The most posterior and largest are the 12 molars, which have several pointed cusps used to crush food so it is ready for swallowing.
did you know
The third members of each set of three molars, top and bottom, are commonly referred to as the wisdom teeth because their eruption is commonly delayed until early adulthood. It is not uncommon for wisdom teeth to fail to erupt; that is, they remain impacted. In these cases, the teeth are typically removed by orthodontic surgery.

Permanent and Deciduous Teeth—This figure of two human dentitions shows the arrangement of teeth in the maxilla and mandible, and the relationship between the deciduous and permanent teeth. Also, the approximate ages when specific teeth erupt are shown.

The teeth are secured in the alveolar processes (sockets) of the maxilla and the mandible.

Gingivae (commonly called the gums) are soft tissues that line the alveolar processes and surround the necks of the teeth. Teeth are also held in their sockets by a connective tissue called the periodontal ligament.

The two main parts of a tooth are the crown, which is the portion projecting above the gum line, and the root, which is embedded within the maxilla and mandible. Both parts contain an inner pulp cavity, containing loose connective tissue through which run nerves and blood vessels. The region of the pulp cavity that runs through the root of the tooth is called the root canal. Surrounding the pulp cavity is dentin, a bone-like tissue. In the root of each tooth, the dentin is covered by an even harder bone-like layer called cementum. In the crown of each tooth, the dentin is covered by an outer layer of enamel, the hardest substance in the body.

IN CONTEXT

Tooth Decay

Although enamel protects the underlying dentin and pulp cavity, it is still nonetheless susceptible to mechanical and chemical erosion, or what is known as tooth decay. The most common form, dental caries (cavities), develop when colonies of bacteria feeding on sugars in the mouth release acids that cause soft tissue inflammation and degradation of the calcium crystals of the enamel.

The Structure of the Tooth—This longitudinal section through a molar in its alveolar socket shows the relationships between enamel, dentin, and pulp.

terms to know
Dentes
Teeth.
Incisor
A midline, chisel-shaped tooth used for cutting into food.
Cuspid (also, Canine)
A pointed tooth used for tearing and shredding food.
Premolar (also, Bicuspid)
A transitional tooth used for mastication, crushing, and grinding food.
Molar
A tooth used for crushing and grinding food.
Gingivae
The gums.
Crown
The portion of tooth visible superior to the gum line.
Root
The portion of a tooth embedded in the alveolar processes beneath the gum line.
Pulp Cavity
The deepest portion of a tooth, containing nerve endings and blood vessels.
Dentin
The bone-like tissue immediately beneath the enamel of the crown or cementum of the root of a tooth.
Cementum
Bone-like tissue covering the root of a tooth.
Enamel
The covering of the dentin of the crown of a tooth.


2. Swallowing

Deglutition is another word for swallowing—the movement of food from the mouth to the stomach. The entire process takes about 4 to 8 seconds for solid or semisolid food, and about 1 second for very soft food and liquids. Although this sounds quick and effortless, deglutition is, in fact, a complex process that involves both the skeletal muscle of the tongue and the muscles of the pharynx and esophagus. It is aided by the presence of mucus and saliva.

There are three stages in deglutition: the voluntary phase, the pharyngeal phase, and the esophageal phase. The autonomic nervous system controls the latter two phases.

Deglutition—Deglutition includes the voluntary phase (on the left) and two involuntary phases: the pharyngeal phase (middle) and the esophageal phase (right).

The voluntary phase of deglutition (also known as the oral or buccal phase) is so-called because you can control when you swallow food. In this phase, chewing has been completed, and swallowing is set in motion. The tongue moves upward and backward against the palate, pushing the bolus to the back of the oral cavity and into the oropharynx. Other muscles keep the mouth closed and prevent food from falling out. At this point, the two involuntary phases of swallowing begin.

The pharyngeal phase is the first step of swallowing that is irreversible. In this phase, there is rapid muscle contraction that forces the bolus into the esophagus. This occurs by stimulation of receptors in the oropharynx that sends impulses to the deglutition center (a collection of neurons that controls swallowing) in the medulla oblongata. Impulses are then sent back to the uvula and soft palate, causing them to move upward and close off the nasopharynx. The laryngeal muscles also constrict to prevent aspiration of food into the trachea. At this point, deglutition apnea takes place, which means that breathing ceases for a very brief time. Contractions of the pharyngeal constrictor muscles move the bolus through the oropharynx and laryngopharynx. Relaxation of the upper esophageal sphincter then allows food to enter the esophagus.

The entry of food into the esophagus marks the beginning of the esophageal phase of deglutition, which is when the bolus moves through the esophagus to the stomach and the initiation of peristalsis. As in the previous phase, the complex neuromuscular actions are controlled by the medulla oblongata. Peristalsis propels the bolus through the esophagus and toward the stomach. The circular muscle layer of the muscularis contracts, pinching the esophageal wall and forcing the bolus forward. At the same time, the longitudinal muscle layer of the muscularis also contracts, shortening this area and pushing out its walls to receive the bolus. In this way, a series of contractions keeps moving food toward the stomach. When the bolus nears the stomach, distention (enlargement) of the esophagus initiates a short reflex relaxation of the lower esophageal sphincter that allows the bolus to pass into the stomach. During the esophageal phase, esophageal glands secrete mucus that lubricates the bolus and minimizes friction.

terms to know
Deglutition
The three-stage process of swallowing.
Voluntary Phase
The initial phase of deglutition in which the bolus moves from the mouth to the oropharynx.
Pharyngeal Phase
The second phase of deglutition in which muscle contraction forces the bolus into the esophagus.
Esophageal Phase
The third phase of deglutition in which the bolus moves through the esophagus to the stomach.


3. Peristalsis

As you're chewing food, your tongue will push the food against your palate, which is also known as the roof of your mouth, and mix it with saliva. Once the mouth is done mechanically and chemically digesting the food, it is then swallowed; the chewed ball of food that is swallowed is called a bolus.

As the food is swallowed, it will move down through your esophagus toward your stomach via peristalsis. As you previously learned, peristalsis is characterized by wave-like contraction of the muscles of the digestive tract that propels food through it. Your esophagus will contract just above the bolus and push it downward. Then, a new contraction will go above it again and push it down farther.

Peristalsis is also how your stomach mechanically breaks the bolus down further and how the intestines push food through your digestive tract.

Peristalsis Moving a Food Bolus

terms to know
Palate
Also known as the "roof" of your mouth; the tongue presses food against the palate and mixes it with saliva to break it down before swallowing.
Bolus
The ball of chewed food that is swallowed.
Peristalsis
Wave-like muscle contractions.

summary
In this lesson, you learned about chewing and swallowing and their role in digestion. Specifically, you looked at the role of the mouth and how the different features of the mouth, such as the teeth, facilitate chewing and the first stages of digestion. Then, you learned about the process of swallowing (deglutition), and that there are three phases of this process: the voluntary phase, the pharyngeal phase, and the esophageal phase. Finally, you learned about the role of peristalsis in moving food down the esophagus towards the stomach.

SOURCE: THIS TUTORIAL HAS BEEN ADAPTED FROM (1) OPENSTAX “ANATOMY AND PHYSIOLOGY 2E”. ACCESS FOR FREE AT OPENSTAX.ORG/BOOKS/ANATOMY-AND-PHYSIOLOGY-2E/PAGES/1-INTRODUCTION. (2) OPENSTAX “MICROBIOLOGY”. ACCESS FOR FREE AT OPENSTAX.ORG/BOOKS/MICROBIOLOGY/PAGES/1-INTRODUCTION. LICENSING (1 & 2): CREATIVE COMMONS ATTRIBUTION 4.0 INTERNATIONAL.

Terms to Know
Bolus

The ball of chewed food that is swallowed.

Cementum

Bone-like tissue covering the root of a tooth.

Crown

The portion of tooth visible superior to the gum line.

Cuspid (also, Canine)

A pointed tooth used for tearing and shredding food.

Deglutition

The three-stage process of swallowing.

Dentes

Teeth.

Dentin

The bone-like tissue immediately beneath the enamel of the crown or cementum of the root of a tooth.

Enamel

The covering of the dentin of the crown of a tooth.

Esophageal Phase

The third phase of deglutition in which the bolus moves through the esophagus to the stomach.

Gingivae

The gums.

Hard Palate

The bony structure that forms the roof of the mouth and floor of the nasal cavity, formed by the palatine process of the maxillary bones and the horizontal plate of the palatine bones.

Incisor

A midline, chisel-shaped tooth used for cutting into food.

Labia

The lips.

Molar

A tooth used for crushing and grinding food.

Oral Cavity

Referred to as the cheeks, tongue, and palate which frame the mouth.

Oral Vestibule

The part of the mouth bounded externally by the cheeks and lips, and internally by the gums and teeth.

Palate

Also known as the "roof" of your mouth; the tongue presses food against the palate and mixes it with saliva to break it down before swallowing.

Peristalsis

Wave-like muscle contractions.

Pharyngeal Phase

The second phase of deglutition in which muscle contraction forces the bolus into the esophagus.

Premolar (also, Bicuspid)

A transitional tooth used for mastication, crushing, and grinding food.

Pulp Cavity

The deepest portion of a tooth, containing nerve endings and blood vessels.

Root

The portion of a tooth embedded in the alveolar processes beneath the gum line.

Soft Palate

The posterior region of the bottom portion of the nasal cavity that consists of skeletal muscle.

Voluntary Phase

The initial phase of deglutition in which the bolus moves from the mouth to the oropharynx.